A number of algorithms have been proposed aimed at tackling the problem of learning "Gene Linkage" within the context of genetic optimisation, that is to say, the problem of learning which groups of co-adapted genes should be inherited together during the recombination process. These may be seen within a wider context as a search for appropriate relations which delineate the search space and "guide" heuristic optimisation, or, alternatively, as a part of a comprehensive body of work into Adaptive Evolutionary Algorithms. In this paper, we consider the learning of Gene Linkage as an emergent property of adaptive recombination operators. This is in contrast to the behaviour observed with fixed recombination strategies in which there is no correspondence between the sets of genes which are inherited together between generations, other than that caused by distributional bias. A discrete mathematical model of Gene Linkage is introduced, and the common families of recombi...